Unfortunately for the animals,the cages were small and impossible to hide in.The zoo environment was anything but natural.(3) ____ the zoo keepers took good care of the animals and fed them well,many of the animals did not thrive;they behaved in strange ways,and they often became ill.
In modern zoos,people can see animals in more natural habitats.The animals are given more (4) ____ in large areas so that they can live more comfortably as they would in nature.Even the appearance of zoos has changed.Trees and grass grow in the cages,and streams of water flow (5) ____ the areas that animals live in.
A.of
B.Although
C.where
D.over
E.freedom
_______